Fruit

The term fruit has different meanings depending on context. In botany, a fruit is the ripened ovary—together with seeds—of a flowering plant. In many species, the fruit incorporates the ripened ovary and surrounding tissues. Fruits are the means by which flowering plants disseminate seeds. In cuisine, when discussing fruit as food, the term usually refers to those plant fruits that are sweet and fleshy, examples of which include plums, apples and oranges. However, a great many common vegetables, as well as nuts and grains, are the fruit of the plant species they come from. No single terminology really fits the enormous variety that is found among plant fruits. The cuisine terminology for fruits is inexact and will remain so. The term false fruit (pseudocarp, accessory fruit) is sometimes applied to a fruit like the fig (a multiple-accessory fruit; see below) or to a plant structure that resembles a fruit but is not derived from a flower or flowers. Some gymnosperms, such as yew, have fleshy arils that resemble
fruits and some junipers have berry-like, fleshy cones. The term "fruit" has also been inaccurately applied to the seed-containing female cones of many conifers.

With most fruits pollination is a vital part of fruit culture, and the lack of knowledge of pollinators and pollenizers can contribute to poor crops or poor quality crops. In a few
species, the fruit may develop in the absence of pollination/fertilization, a process known as parthenocarpy. Such fruits are seedless. A plant that does not produce fruit is known as acarpous, meaning "without fruit"

Botanic Fruit and Culinary Fruit

Many foods are botanically fruit but are treated as vegetables in cooking. These include cucurbits (e.g., squash, pumpkin, and cucumber), tomato, peas, beans, corn,
eggplant (aubergine), and sweet pepper, spices, such as allspice and chillies. Occasionally, though rarely, a culinary "fruit" will not be a true fruit in the botanical sense. For example, rhubarb may be considered a fruit, though only the astringent petiole is edible. In the commercial world, European Union rules define carrot as a fruit for the purposes of measuring the proportion of "fruit" contained in carrot jam. In the culinary sense, a fruit is usually any sweet tasting plant product associated with seed(s), a vegetable is any savoury or less sweet plant product, and a nut any hard, oily, and shelled plant product.

Although a nut is a type of fruit, it is also a popular term for edible seeds, such as peanut (which is actually a legume), pistachio and walnut. Technically, a cereal grain is a fruit termed a caryopsis. However, the fruit wall is very thin and fused to the seed coat so almost all of the edible grain is actually a seed. Therefore, cereal grains, such as corn, wheat and rice are better considered edible seeds, although some references list them as fruits. Edible gymnosperms seeds are often misleadingly given fruit names, e.g. pine nuts, ginkgo nuts, and juniper berries.

Fruit Development

A fruit is a ripened ovary. After the ovule in an ovary is fertilized in a process known as pollination, the ovary begins to ripen. The ovule develops into a seed and the ovary wall pericarp may become fleshy (as in berries or drupes), or form a hard outer covering (as in nuts). In some cases, the sepals, petals and/or stamens and style of the flower fall off. Fruit development continues until the seeds have matured. With some multiseeded fruits the extent to which the flesh develops is proportional to the number of
fertilized ovules.

The wall of the fruit, developed from the ovary wall of the flower, is called the pericarp. The pericarp is often differentiated into two or three distinct layers called the exocarp
(outer layer - also called epicarp), mesocarp (middle layer), and endocarp (inner layer). In some fruits, especially simple fruits derived from an inferior ovary, other parts of the flower (such as the floral tube, including the petals, sepals, and stamens), fuse with the ovary and ripen with it. The plant hormone ethylene causes ripening. When such other floral parts are a significant part of the fruit, it is called an accessory fruit. Since other parts of the flower may contribute to the structure of the fruit, it is important to study flower structure to understand how a particular fruit forms.

Fruits are so varied in form and development, that it is difficult to devise a classification scheme that includes all known fruits. Many common terms for seeds and fruit are incorrectly applied, a fact that complicates understanding of the terminology. Seeds are ripened ovules; fruits are the ripened ovaries or carpels that contain the seeds. To these two basic definitions can be added the clarification that in botanical terminology, a nut is a type of fruit and not another term for seed.

There are three basic types of fruits:

Simple fruit
Aggregate fruit
Multiple fruit

Simple Fruit

Simple fruits can be either dry or fleshy and result from the ripening of a simple or compound ovary with only one pistil. Dry fruits may be either dehiscent (opening to discharge seeds), or indehiscent (not opening to discharge seeds).

Types of dry, simple fruits (with examples) are:
achene - (buttercup)
capsule - (Brazil nut)
caryopsis - (wheat)
fibrous drupe - (coconut, walnut)
follicle - (milkweed)
legume - (pea, bean, peanut)
loment nut - (hazelnut, beech, oak acorn)
samara - (elm, ash, maple key)
schizocarp - (carrot)
silique - (radish)
silicle - (shepherd's purse)
utricle - (beet)
Fruits in which part or all of the pericarp (fruit wall) is fleshy at maturity are simple fleshy fruits.

Types of fleshy, simple fruits (with examples) are:
berry - (tomato, avocado)
stone fruit or drupe (plum, cherry, peach, olive)
false berry - accessory fruits (banana, cranberry)
pome - accessory fruits (apple, pear, rosehip)

Aggregate fruit

An aggregate fruit, or etaerio, develops from a flower with numerous simple pistils. An example is the raspberry, whose simple fruits are termed drupelets because each
is like a small drupe attached to the receptacle. In some bramble fruits (such as blackberry) the receptacle is elongated and part of the ripe fruit, making the blackberry an
aggregate-accessory fruit. The strawberry is also an aggregate-accessory fruit, only one in which the seeds are contained in achenes. In all these examples, the fruit develops from a single flower with numerous pistils.

Multiple Fruit

A multiple fruit is one formed from a cluster of flowers (called an inflorescence). Each flower produces a fruit, but these mature into a single mass. Examples are the pineapple, edible fig, mulberry, osage-orange, and breadfruit.In the photograph on the right, stages of flowering and fruit development in the noni or Indian mulberry (Morinda citrifolia) can be observed on a single branch. First an inflorescence of white flowers called a head is produced. After fertilization, each flower develops into a drupe, and as the drupes expand, they become connate (merge) into a multiple fleshy fruit called a syncarpet.

There are also many dry multiple fruits, e.g.
Tuliptree, multiple of samaras.
Sweet gum, multiple of capsules.
Sycamore and teasel, multiple of achenes.
Magnolia, multiple of follicles.

Seedless fruits

Seedlessness is an important feature of some fruits of commerce. Commercial cultivars of bananas and pineapples are examples of seedless fruits. Some cultivars of citrus
fruits (especially navel oranges and mandarin oranges), table grapes, grapefruit, and watermelons are valued for their seedlessness. In some species, seedlessness is the result of parthenocarpy, where fruits set without fertilization. Parthenocarpic fruit set may or may not require pollination. Most seedless citrus fruits require a pollination stimulus; bananas and pineapples do not. Seedlessness in table grapes results from the abortion of the embryonic plant that is produced by fertilization, a phenomenon known as stenospermocarpy which requires normal pollination and fertilization.